Your engine has TWO temperature sensors.

The single-blade sensor, under intake header #2, is for the temp gauge (only).

The double-blade sensor, under intake header #3, is for the FI ecu. It tells the fuel injection system whether your engine is hot, cold, or frigid, and the FI systems adjusts the fuel mixture accordingly.

The resistance of the sensor varies inversly with the temperature. A hot sensor (high temp) means a lower resistance. A cold sensor means high resistance.

If a wire broke at the push-on connector, the the FI ecu would measure infinitely high resistance, which it would interpret as a very cold engine. The ecu would pour in the fuel. On the other hand, if the wires were shorted or the sensor had an internal short, the resistance would be low and the ecu would think the engine was hot -- and reduce the fuel accordingly.

"Could it be something other than the sensors causing this problem?"

It could be a bad (or failing) fuel pressure regulator or a bad (or failing) AMM. I wouldn't expect your error code for either of these problems, however.
--
Don Foster (near Cape Cod, MA)